For the model (5.3), what is the expected value of the within-lineage mean square at time (t)

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For the model (5.3), what is the expected value of the within-lineage mean square at time \(t\) ? For the Brownian-motion model (5.4), show that the variance of \(Y_{u}\) increases linearly with time. What is the expected value of the within-lineage mean square?

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