What Big Five Personality Trait was found to have a negative relationship to prejudice? (A negative relationship between two variables means that the higher amount of the first variable means a lower amount of the second variable. In this case the higher amount of of the Big Five Personality trait means that there is a lower amount of prejudice) How do you explain this finding?

Article: What Matters Most to Prejudice: Big Five Personality, Social Dominance Orientation, or Right-Wing Authoritarianism?

Please include references.