Develop a word equation relating the population of Atlantic Puffins Nt in one year to the population Nt+1 in the next year.

The word equation might involve things like

Number of Atlantic Puffins born in the wild during the year

Atlantic Puffins removed for captive breeding during the year

Atlantic Puffins reintroduced to the wild from captive breeding during the year

Atlantic Puffins killed by predators during the year

Atlantic Puffin deaths from disease during the year