Consider the separation of a mixture containing the following protein-analytes: ovalbumin (molecular weight ~ 43,000 Da), glutamate dehydrogenase (molecular weight ~ 290,000 Da), serum albumin (molecular weight ~ 67,000 DA), lactate dehydrogenase (molecular weight ~ 140,000 Da), and cytochrome c (molecular weight ~ 12,400) on a size-exclusion column with a fractionation range of 15,000 - 150,000 Da. In what order would the proteins elute from the column and why?